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To clarify, those wetlands etc. are being treated as holes in the nature_reserve (wrong), as evidenced by their being drawn with a nature_reserve dashed border and without the ducky nature_reserve fill pattern.  This is in addition to the wetlands etc. also being treated as holes in the wood (correct).  I say wrong/correct because they're only marked as inners of the wood multipolygon, not of  the nature_reserve, though both share the same outer way.

I'm not familiar with osmarender, but the rendering seems correct on all the slippy renderers I have tried (Mapnik, OCM, MapQuest, Humanitarian, ...), and JOSM had no complaints when validating.  So I *think* it's a legal construct.  (EDIT: On closer inspection, it seems OCM and Humanitarian get it "wrong", but it's extremely hard to notice, 1-pixel borders.  Mapnik does it "right", because wrong would be very noticeable.)  Maybe I should just duplicate the outer way...

Anyway, thanks for the quick response.  I doubt it's a theme bug; I simply mentioned it here for lack of knowing where to direct a bug report, and in any case, it's probably a minor one.  Cheers.

A beautiful theme, thank you very much!  (Voluntary UK, in Locus Pro, Andromaps)

I think I've found an obscure bug, but also a workaround.  I'm seeing lakes, wetlands, meadows, etc. being drawn with a dashed border, which is unattractive and apparently not intended.  These are inner members of a wood multipolygon, the outer way of which (307333650) is also the outer way of a nature_reserve multipolygon.  (I think this is legal.)

When I commented out the two "line" elements in the XML file, the ones following the nature_reserve keyword (line 808), the problem went away.  So apparently, the boundary decoration of the nature_reserve "jumped over and down" to the interior holes of its cousin, the wood multipolygon.  Strange but apparently true.

I've noticed this behavior in some of the other themes too.  Anyway, I have a workaround, so I'm quite happy.